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Den 16. film fra Pixar udspiller sig i en alternativ fortid, hvor Jordens dinosaurer ikke blev udryddet af et gigantisk meteornedslag. I stedet lever de store øgler side om side med menneskene. Apatosaurusungen Arlo bliver forældreløs, da han mister sin familie i en flodbølge. I forsøget på at finde hjem møder Arlo menneskeungen Tot. Venskabet med den vilde, primitive dreng er med til at modne den frygtsomme dinosaurus.
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