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Historia sobre una guerra extraterrestre que tiene lugar en la Tierra. Dos razas de robots transformables -los villanos decepticons y los amistosos autobots- llegan a nuestro planeta en busca de la "chispa vital", o "el cubo", una misteriosa fuente de poder. Aquí entablarán una disputa con los humanos de por medio, que tomarán partido por los autobots. La Tierra es la salida para estas máquinas, pues rica en minerales y químicos, representa la salvación para los autómatas, quienes emprenderán una terrible guerra por el preciado Cubo. Y, en medio de todo, un espabilado adolescente, Sam Witwicky (Shia LaBeouf), que sólo pretende conquistar a la bella Mikaela (Megan Fox), se convierte en la clave de una guerra que puede destruir a la humanidad.
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