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1800-luvun Englannissa Charles Darwinin elämänhalu sammuu, kun hänen vanhin tyttärensä kuolee. Masentunut Darwin vetäytyy työhuoneensa uumeniin kehittelemään teoriaa, joka voi muuttaa maailmaa mutta on samalla kehittäjälleen vaarallinen, sillä se tekee hänestä uskonnollisten kiihkoilijoiden raivon kohteen.
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