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圣诞节临近,狮子亚历克斯(本·斯蒂勒 Ben Stiller 配音)、斑马马蒂(克里斯·洛克 Chris Rock 配音)、长颈鹿迈尔曼(大卫·休默 David Schwimmer 配音)以及河马格洛里亚(贾达·萍克·史密斯 Jada Pinkett Smith 配音)决定正式告别马达加斯加,回到久违的纽约共度佳节。冷冷清清的送别晚会没能熄灭它们归去的热情,好伙伴们燃起热气球正式出发,可是却被狐猴王朱利安当作红色妖盗击落下来。不久,真的红色妖盗出现夜空,混乱之中亚历克斯阴差阳错将其打落,结果却发现所谓的红色妖盗竟然是圣诞老人!倒霉的是圣诞老人被砸失忆,为了不让全世界小朋友失望,亚历克斯他们和企鹅帮决定充当圣诞老人的助手,将礼物送到世界每一个角落。
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